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Swarming generates rebel workers in honeybees

Workers which develop in a temporarily queenless colony, for example during swarming, have more ovarioles, less developed hypopharyngeal glands and more developed mandibular glands. This can be explained by reduced relatedness of the workers to the new queen produced during swarming Swarming generates rebel workers in honeybees,
Current Biology
, Volume 22, p.707-711, (2012)
[7].

See also:
http://www.sciencedirect.com/science/article/pii/S0960982212002758